List of Asian animals extinct in the Holocene
This list of Asian animals extinct in the Holocene features animals known to have become extinct in the last 12,000 years on the Asian continent and its islands.
Many extinction dates are unknown due to a lack of relevant information.
Species from Christmas Island, an outlying island of Australia, are listed in List of Australia-New Guinea species extinct in the Holocene.
Mammals
Undated
Common name scientific name |
Range |
---|---|
Flores cave rat Spelaeomys florensis |
Flores, Indonesia[1] |
Ochotona transcaucasica | Caucasus[2] |
Prehistoric
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Panay golden-crowned flying foxAcerodon jubatus lucifer | Extinct subspecies of the giant golden-crowned flying fox (Acerodon jubatus). | ||
Bubal hartebeest Alcelaphus buselaphus buselaphus |
1200-586 BCE[3] | North Africa and southern Levant | |
Alor Island giant rat Alormys aplini |
1050 BCE[4] | Alor Island, Indonesia | |
Steppe bison Bison priscus |
2550-2450 BCE[5] | Northern Eurasia and North America | |
Indian aurochs Bos primigenius namadicus |
1800 BCE[6] | Indian subcontinent | |
Eurasian aurochs Bos primigenius primigenius |
1900-1745 BCE[7] | Mid-latitude Eurasia[8] | |
Cebu tamaraw Bubalus cebuensis |
Pleistocene or Holocene | The Cebu tamaraw stood only 75 cm (2 ft 6 in) and weighed about 150 to 160 kg (330 to 350 lb).[9][10] The species was smaller than the modern Mindoro tamaraw. | |
Short-horned water buffalo Bubalus mephistopheles |
1750-1650 BCE[11] | South, central, and east China | |
Carpomys dakal | 0 CE | Philippines
A giant cloud rat with size rivaling the Phloeomids reaching around 2 kg. In several Philippine languages, dakal means big or large.[12] |
|
Crateromys ballik | 0 CE | Philippines
Another murine with fossils found in Cagayan, regardless of its large size it was named ballik after the Dupaningan Agta word for small. This is in reference for its relatively smaller size than the extant cloud runners of the same species (i.e. giant bushy-tailed cloud rat).[13] |
|
Miyako roe deer Capreolus tokunagai |
9050-8050 BCE[4] | Miyako Island, Ryukyu, Japan | |
Woolly rhinoceros Coelodonta antiquitatis |
8050-7650 BCE[14] | Northern Eurasia | |
Buhler's coryphomys Coryphomys buehleri |
50 BCE[4] | Timor | |
Timor giant rat Coryphomys musseri |
50 BCE[4] | Timor | |
Chinese cave hyena Crocuta crocuta ultima |
9550 BCE[15] | East Asia | |
Miyako long-tailed rat Diplothrix miyakoensis |
9050-8050 BCE[4] | Miyako Island, Ryukyu, Japan | |
Syrian elephant Elephas maximus asurus |
800-700 BCE[16] | Mesopotamia | |
European wild ass Equus hemionus hydruntinus |
1294-1035 BCE[17] | Europe and southwest Asia | |
Lena horse Equus (ferus) lenensis[18] |
400-1 BCE[19] | Northern Siberia | |
Equus ovodovi | c. 1500 BCE[20] | Southern Siberia to Northern China | |
Cypriot genet Genetta plesictoides |
9050 BCE[4] | Cyprus | |
Hooijer's giant rat Hooijeromys nusantenggara |
1050 BCE[4] | Lesser Sunda Islands, Indonesia | |
Imperial gibbon Junzi imperialis |
c. 240 BCE[21] | Shaanxi, China? | |
Woolly mammoth Mammuthus primigenius |
1795-1675 BCE[22] | Northern Eurasia and North America | |
Irish elk Megaloceros giganteus |
4901-4831 BCE[23][19] | Europe and southern Siberia | |
Asian straight-tusked elephant Palaeoloxodon namadicus |
7330-6250 BCE (unconfirmed)[24] |
South and east Asia | |
Verhoeven's giant tree rat Papagomys theodorverhoeveni |
1050 BCE[4] | Flores, Indonesia | |
Sumba Island giant rat Raksasamys tikusbesar |
1935-1700 BCE[4] | Sumba Island, Indonesia |
Recent
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Caucasian moose Alces alces caucasicus |
c. 1900[25] | Northern Caucasus and Transcaucasian coast of the Black Sea | |
Caucasian wisent Bison bonasus caucasicus |
1927[26] | Caucasus Mountains | |
Hokkaidō wolf Canis lupus hattai |
c. 1889[27] | Hokkaidō, Sakhalin, Kamchatka, Iturup and Kunashir[28] | |
Japanese wolf Canis lupus hodophilax |
1905[27] | Honshū, Shikoku, and Kyūshū, Japan | |
Syrian wild ass Equus hemionus hemippus |
1927[29] | Fertile Crescent | |
Queen of Sheba's gazelle Gazella bilkis |
1951[30] | Southwestern Yemen | |
Saudi gazelle Gazella saudiya |
1970[31] | Arabian Peninsula | |
Steller's sea cow Hydrodamalis gigas |
before 1768[32] | Bering Sea | |
Hokkaidō otter Lutra lutra whiteleyi |
1950s[33] | Hokkaidō and southern Kuril Islands[34] | |
Japanese otter Lutra nippon |
1983[34] | Honshū, Shikoku, and Kyūshū, Japan | |
Formosan clouded leopard Neofelis nebulosa brachyura |
1983[35] | Taiwan | |
Bali tiger Panthera tigris balica |
1937[36] | Bali, Indonesia | |
Javan tiger Panthera tigris sondaica |
1984[36] | Java, Indonesia | |
Caspian tiger Panthera tigris virgata |
1979[36] | Western and Central Asia | |
Sturdee's pipistrelle Pipistrellus sturdeei |
1889[37] | Haha-jima, Bonin Islands, Japan | |
Vietnamese rhinoceros Rhinoceros sondaicus annamiticus |
2010[38] | Vietnam, Laos, Cambodia, and eastern Thailand | |
Indian Javan rhinoceros Rhinoceros sondaicus inermis |
1920[38] | Northeastern India, Bangladesh, and Myanmar | |
Schomburgk's deer Rucervus schomburgki |
1938[39] | Central Thailand | |
Japanese sea lion Zalophus japonicus |
1951[40] | Japanese archipelago and Korea |
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Kouprey Bos sauveli |
1969[41] | Notheastern Cambodia | |
Jolo sika deerCervus nippon soloensis | Introduced during the Holocene (unknown date). No current population census and may be possibly extinct in Sulu.[42] Although it is still listed by the IUCN as extant resident introduced population.[43] | Jolo island, Southern Phillippines (anciently introduced) | |
Ilin Island cloudrunner Crateromys paulus |
1953[44] | Mindoro and Ilin Island, Philippines
Locally known as siyang,[45] it is known from one specimen collected in 1953, and has never been formally studied in the wild.[46] |
|
Northern Sumatran rhinoceros Dicerorhinus sumatrensis lasiotis |
unknown[47] | India, Bhutan, Bangladesh, and Myanmar | |
Baiji Lipotes vexillifer |
2002[48] | Middle and lower Yangtze River, China | |
Gloomy tube-nosed bat Murina tenebrosa |
1962[49] | Tsushima and possibly Yaku Island, Japan | |
Cebu warty pig Sus cebifrons cebifrons |
1960s[50] | Cebu, Philippines[51]
The Cebu warty pig previously lived on the island of Cebu before becoming extinct in modern times, primarily due to habitat destruction and human exploitation. The subspecies was declared extinct in 2000, but other warty pig subspecies still survive on other Philippine islands. |
|
Malabar large-spotted civet Viverra civettina |
1989[52] | Western Ghats, India |
Local
Common name scientific name |
Extinction date | Range | Reintroduction | Image |
---|---|---|---|---|
Lowland wisent Bison bonasus bonasus |
18th century | Central Europe to southern Siberia | 1946 (Caucasus) 1982 (Altai)[53] |
|
Bactrian camel Camelus bactrianus |
after 4000-3000 BCE[A] | Central and eastern Asian steppe[55] | ||
Dromedary Camelus dromedarius |
404 BCE[56] | Arabian Peninsula | ||
Dhole Cuon alpinus |
5,000 BCE[57] | Locally extinct in Palawan, Philippines
Lived on Palawan during the Pleistocene and Early Holocene.[58] With fossil remains similar to Canis familiaris, it remains inconclusive. But the presence of a wild dog crossing from the Sunda shelf to Palawan is cited to be more likely than an early arrival of a domesticated dog.[59][60] |
||
Pere David's deer Elaphurus davidianus |
c. 400[61] | Northeastern China | 1985[62] | |
African wild ass Equus africanus |
3000 BCE[63] | North Africa, Levant, and Arabian Peninsula | ||
Przewalski's horse Equus ferus przewalskii |
1969 | Central and eastern Asian steppe | 1994[64] | |
Lesser ranee mouseHaeromys pusillus | Possibly extirpated in Calauit islands and Palawan.[65] | |||
Hippopotamus Hippopotamus amphibius |
1200-586 BCE[3] | Subsaharan Africa, Egypt, and the Levantine coast[66] | ||
Arabian oryx Oryx leucoryx |
1972[67] | Arabian Peninsula | 1982[68] | |
Muskox Ovibos moschatus |
615-555 BCE[19] | Northern Eurasia and North America | 1974[69] | |
South China tiger Panthera tigris amoyensis |
c. 2000[70][71] | Southern China | ||
Tiger Panthera tigris ssp. |
5,000 BCE[72] | Locally extinct in Palawan, Philippines
Two articulated phalanges and another phalanx piece were found amidst an assemblage of other animal bones and stone tools in Ille Cave near the village of New Ibajay. From the stone tools, besides the evidence for cuts on the bones, and the use of fire, it would appear that early humans had accumulated the bones.[73] Additionally, the condition of the tiger subfossils, dated to approximately 12,000 to 9,000 years ago, differed from other fossils in the assemblage, dated to the Upper Paleolithic. The tiger subfossils showed longitudinal fracture of the cortical bone due to weathering, which suggests that they had post-mortem been exposed to light and air. Tiger parts were commonly used as amulets in South and Southeast Asia, so it may be that the tiger parts were imported from elsewhere, as is the case with tiger canine teeth, which were found in Ambangan sites dating to the 10th to 12th centuries in Butuan, Mindanao. On the other hand, the proximity of Borneo and Palawan also makes it likely that the tiger had colonized Palawan from Borneo before the Early Holocene.[74][75] |
||
Bornean bearded pigSus barbatus | Reports of Bornean pigs on the islands of the Sulu archipelago have been documented, probably driven by their periodically sea crossing habits to reach nearby islands.[76] It is classified by the IUCN Red List as possibly extinct in the area.[77] | Locally extinct in the Sulu archipelago, Phillippines | ||
Lesser kudu Tragelaphus imberbis |
1968?[78][63] | East Africa and western Arabian Peninsula |
Birds
Undated
Common name scientific name |
Range |
---|---|
Daito bush warbler Horornis diphone restricta |
Daito Islands, Japan[79] |
Prehistoric
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Bennu heron Ardea bennuides |
2550 BCE[80] | Arabian Peninsula | |
East Asian ostrich Struthio anderssoni |
6950 BCE[81] | Northern China and Mongolia | |
Asian ostrich Struthio asiaticus |
7600-6245 BCE[82] | Eastern Europe through Kazakhstan to India and China[83] |
Recent
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Car Nicobar sparrowhawk Accipiter butleri butleri |
1901[84] | Car Nicobar, Nicobar Islands | |
Iwo Jima rail Amaurornis cinerea brevipes |
1911[84] | Naka Iwo Jima and Minami Iwo Jima, Bonin Islands | |
Luzon sarus craneAntigone antigone luzonica | 1960s | Species extirpated in the Philippines, subspecies became extinct in the 1960s.[85] No distinctive characteristic was observed or studied in the Philippine population compared to other subspecies.[86] | |
Bonin grosbeak Carpodacus ferreorostris |
1828[87] | Bonin Islands, Japan | |
Cyprus dipper Cinclus cinclus olympicus |
1945[88] | Cyprus | |
Bonin wood pigeon Columba versicolor |
1889[89] | Bonin Islands, Japan | |
Ryukyu wood pigeon Columba jouyi |
1936[84] | Ryukyu, Japan | |
Cebu bar-bellied cuckooshrikeCoracina striata cebuensis | Extinct in the 20th century.[90] | ||
Cebu hanging parrot Loriculus philippensis chrysonotus |
1943[84] | Cebu, Philippines
A recognized subspecies of the Philippine hanging parrot, generally believed to be extinct.[91] |
|
Siquijor hanging parrot Loriculus philippensis siquijorensis |
1908[84] | Siquijor, Philippines
Like the Cebu subspecies, this subspecies is also regarded as extinct.[92] |
|
Bonin nankeen night heron Nycticorax caledonicus crassirostris |
1889[84] | Chichi-jima and Nakōdo-jima, Bonin Islands | |
Cebu dark-throated orioleOriolus steerii assimilis | 1906 | Initially considered a separate species by Tweede in 1878.[93] Last sighted in 1906. | |
Ticao tarictic hornbill Penelopidis panini ticaensis |
1971[84] | Ticao Island, Philippines
A subspecies of the Visayan tarictic hornbill that lived in Ticao island. It was declared extinct by the year 2013[94] and last seen in 1971. |
|
Samar bay owlPhodilus badius riverae | 1940s | Extinct on the mid 20th century, with its taxon's validity debated, and was only known from one specimen which was lost in 1945.[95] Bay owls of other subspecies are still considered as rare/accidental species within the region. | |
Bornean Baillon's crake Porzana pusilla mira |
1912[84] | Borneo | |
Daito varied tit Sittiparus varius orii |
1938[84] | Kitadaitōjima, Okinawa, Japan | |
Arabian ostrich Struthio camelus syriacus |
c. 1941[96] | Arabian Peninsula and Near East | |
Tawi-tawi buttonquail Turnix sylvaticus suluensis |
1950s[84] | Jolo and Tawi-tawi, Sulu archipelago, Philippines | |
Spectacled cormorant Urile perspicillatus |
1850[97] | Commander Islands, Russia | |
Bonin thrush Zoothera terrestris |
1828[98] | Bonin Islands, Japan |
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Mukojima white-eye Apalopteron familiare familiare |
1930s[99] | Mukojima, Bonin Islands | |
Sangihe dwarf kingfisher Ceyx fallax sangirensis |
1997[84] | Sangihe Islands, Indonesia | |
Cebu blackish cuckooshrikeCoracina coerulescens altera | Last confirmed sighting in 1906.[100] Unconfirmed sighting in 2000.[101] | ||
Rück's blue flycatcher Cyornis ruckii |
1918[102] | Northern Sumatra coast, Indonesia | |
Cebu white-bellied woodpeckerDryocopus javensis cebuensis | With unconfirmed possible sightings in 1998 and 2000.[103] It is likely critically endangered if not extinct. | ||
White-eyed river martin Eurochelidon sirintarae |
1978[104] | Central Thailand | |
Catanduanes bleeding heartGallicolumba luzonica rubiventris | Based only from one single specimen collected in 1971, lives exclusively in Catanduanes. | ||
Sulu bleeding-heart Gallicolumba menagei |
1891[84] | Tawi-tawi, Sulu archipelago, Philippines
Expedition on 2009, failed to find signs of this species' continued existence.[105] The Sulu bleeding-heart if not extinct, may have been critically endangered and may probably be less than 50 individuals. Other reported sightings may be attributed to other local bleeding-heart species, e.g. Luzon and Mindanao bleeding-hearts. |
|
Negros celestial monarchHypothymis coelestis rabori | Found on Sibuyan and Negros. This subspecies has not been recorded since 1959 and may now be extinct.[106][107] | ||
Slender-billed curlew Numenius tenuirostris |
2001[84] | Western Eurasia and North Africa | |
Himalayan quail Ophrysia superciliosa |
1876[84] | Uttarakhand, India | |
Siau scops owl Otus siaoensis |
1866[84] | Siau Island, Indonesia | |
Spot-billed pelicanPelicanus philippensis | 1960s | Named taxonomically because of abundance in the Philippines up until the 1900s[108] but declined and become extirpated in the country around the 1960s.[109] | |
Cebu brown dovePhapitreron amethystina frontalis | Darker overall appearance compared to the other subspecies of the amethyst brown dove race. It lacks the white the stripe under eye. It is considered as probably extinct. Not seen since 1892. Possible sighting was two individuals in the Alcoy forest in 2004 but needs further proper identification.[110] | ||
Negros fruit dove Ptilinopus arcanus |
1953[111] | Negros Island, Philippines
Originally described in 1953,[112] with an alleged sighting by a local hunter in the 1990s. Currently listed as critically endangered if not extinct. |
|
Pink-headed duck Rhodonessa caryophyllacea |
1949[113] | Northeast India, Bangladesh, and northern Myanmar | |
Crested shelduck Tadorna cristata |
1964[114] | Primorye, Hokkaidō, and South Korea; possibly North Korea and northeastern China | |
Luzon blue-backed parrotTanygnathus everetti duponti | No sightings since the 1970s in the island of Luzon, likely extinct.[115] | ||
Polillo blue-backed parrotTanygnathus everetti freeri | No sightings since 2004 in the island of Polillo, likely extinct.[115] | ||
Javan lapwing Vanellus macropterus |
1940[116] | Java, Indonesia |
Local
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Asian woolly-necked storkCiconia episcopus | Locally extinct in the Philippines.
Last documented in 2007 in Bislig, Surigao del Sur. It is listed by the IUCN as possibly extinct.[117] It was formerly widespread across the islands. |
||
Northern bald ibis Geronticus eremita |
2014[118] | Mediterranean region |
Reptiles
Common name scientific name |
Extinction date | Range |
---|---|---|
Ryukyu tortoise Manouria oyamai |
c. 9050 BCE[119] | Ryukyu, Japan |
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Chinese gharial Hanyusuchus sinensis |
1292-1630[120] | South China and Hainan |
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Nile crocodile Crocodylus niloticus |
c. 1900[121] | Subsaharan Africa, Egypt, and the Levant |
Amphibians
Fish
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Beyşehir bleak Alburnus akili |
1955[141] | Lake Beyşehir, Turkey | |
Anabarilius macrolepis | 1981[142] | Yilong Lake, Yunnan, China | |
Pait Barbodes amarus |
1982[143] | Lake Lanao, Philippines
At least 15 species of endemic cyprinid from Lake Lanao were confirmed to be extinct.[144][145][146] |
|
Baolan Barbodes baoulan |
1991[147] | Lake Lanao, Philippines | |
Bagangan Barbodes clemensi |
1975[148] | Lake Lanao, Philippines | |
Barbodes disa | 1964[149] | Lake Lanao, Philippines | |
Katapa-tapa Barbodes flavifuscus |
1964[150] | Lake Lanao, Philippines | |
Barbodes herrei | 1974[151] | Lake Lanao, Philippines | |
Barbodes katolo | 1977[152] | Lake Lanao, Philippines | |
Kandar Barbodes lanaoensis |
1964[153] | Lake Lanao, Philippines | |
Barbodes manalak | 1977[154] | Lake Lanao, Philippines | |
Bitungu Barbodes pachycheilus |
1964[155] | Lake Lanao, Philippines | |
Bagangan Barbodes palaemophagus |
1975[156] | Lake Lanao, Philippines | |
Barbodes palata | 1964[157] | Lake Lanao, Philippines | |
Bagangan Barbodes resimus |
1964[158] | Lake Lanao, Philippines | |
Barbodes tras | 1976[159] | Lake Lanao, Philippines | |
Bitungu Barbodes truncatulus |
1973[160] | Lake Lanao, Philippines | |
Lost shark Carcharinus obsoletus |
1934[161] | Southern South China Sea | |
Yilong carp Cyprinus yilongensis |
1981[162] | Yilong Lake, Yunnan, China | |
Hula bream Mirogrex hulensis |
1975[163] | Lake Hula, Israel | |
Siamese flat-barbelled catfish Platytropius siamensis |
1975-1977[164] | Chao Phraya and Bang Pakong River basins, Thailand | |
Chinese paddlefish Psephurus gladius |
2003[165] | Yangtze and Yellow River basins, China | |
Syr Darya sturgeon Pseudoscaphirhynchus fedtschenkoi |
1960s[166] | Syr Darya River | |
Schizothorax saltans | 1953[167] | Talas River basin, Kazakhstan | |
Tristramella intermedia | 1970s[168] | Lake Hula, Israel | |
Tristramella magdelainae | 1950s[169] | Damascus, Syria |
Common name scientific name |
Extinction date | Range | Image |
---|---|---|---|
Yangtze sturgeon Acipenser dabryanus |
2000[170] | Yangtze River basin, China | |
Barbodes cataractae | The Phillippines | Additional three species are considered possibly extinct: Barbodes cataractae, B. lindog, and B. sirang. B. lindog and B. sirang have reported sightings within the last ten years (2008 for the former and 2016 for the latter). The B. cataractae on the other hand has not been recorded during the market surveys of 1973 to 2017.[171][172][173] | |
Barbodes lindog | Lake Lanao, Philippines | ||
Barbodes sirang | Lake Lanao, Philippines | ||
Manila bay herringClupea manulensis | Since its description in 1822, there have been no sightings nor documentations that assesses this species' presence.[174] | ||
BiaExyrias volcanus | Endemic to Taal lake, being the only freshwater species within the genus and only located on a limited area; it has not been sighted in surveys since 1996.[175] | ||
Neostethus ctenophorus | Endemic in the Laguna de Bay, it is threatened by industrialization and invasive species. Despite numerous fish surveys it was last seen in 1937.[176] | ||
Kunimasu Oncorhynchus kawamurae |
1940[177] | Lake Tazawa, Japan | |
Rivulated parrotfishScarus rivulatus | Reported extinct in the Philippines by the IUCN during its assessment in 2009 (published in 2012), experiencing massive population reduction by 60-70% in a span of 20–30 years.[178] It, however, was still observed in Siquijor in 2020 comprising 18.6% out of 209 of the collected sample of juvenile parrotfish species.[179] Otherwise, its global population is still under least concern category.[178] | ||
Uling gobySicyopus cebuensis | Locally known as tughud in cebuano. It is classified as data deficient in IUCN Red List but is considered to be possibly extinct in 2015 as the river it resides is polluted.[180] There were alleged sightings in 2019.[181] | ||
Yellow-bellied gobySilhouettea flavoventris | Endemic to Taal lake, described in 1927 with 37 specimens ranging 25–37 mm in length. It has not been sighted since the 1990's.[182] | ||
Beloribitsa Stenodus leucichthys |
1960s[183] | Caspian Sea, Volga, Ural, and Terek River drainages |
Insects
Common name scientific name |
Extinction date | Range |
---|---|---|
Laguna white-legged damselflyRisiocnemis laguna | Has not been rediscovered since its discovery in 1916, and is only known from specimens found in Paete, laguna. With Paete's rapid urbanization and being a heavily populated location with heavy forest degradation. The population of this damselfly is expected to be critically endangered if not extinct.[184] | The Phillippines |
Earthworms
Common name scientific name |
Extinction date | Range |
---|---|---|
Amynthas japonicus | 1820s[185] | Nagasaki?, Japan |
Molluscs
Common name scientific name |
Range | Image |
---|---|---|
Gastrocopta chichijimana | Bonin Islands, Japan[186] | |
Gastrocopta ogasawarana | Bonin Islands, Japan[187] | |
Hirasea planulata | Bonin Islands, Japan[188] | |
Lamellidea monodonta | Bonin Islands, Japan[189] | |
Lamellidea nakadai | Bonin Islands, Japan[190] | |
Littoraria flammea | China[191] | |
Trochoidea picardi | Israel[192] | |
Vitrinula chaunax | Bonin Islands, Japan[193] | |
Vitrinula chichijimana | Bonin Islands, Japan[194] | |
Vitrinula hahajimana | Bonin Islands, Japan[195] |
See also
- Holocene extinction
- List of extinct animals
- List of extinct birds
- Extinct in the wild
- Lazarus taxon
- List of extinct animals of India
- List of extinct animals of the Philippines
Notes
- ^ The extant wild Bactrian camel (C. ferus) from East Turkestan is not the ancestor of C. bactrianus.[54]
References
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