Late Miocene, Botchi Formation, Botchi River, Khabarovsk Krai, Russia
Palaeontological Institute, Russian Academy of Science, Moscow, Russia; specimen #2054/229
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Dehon M, Engel MS, Gérard M, Aytekin AM, Ghisbain G, Williams PH, Rasmont P, Michez D (2019). "Morphometric analysis of fossil bumble bees (Hymenoptera, Apidae, Bombini) reveals their taxonomic affinities". ZooKeys891: 71-118. DOI:10.3897/zookeys.891.36027.
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