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During the Campanian age, a radiation among dinosaur species occurred. In North America, for example, the number of known dinosaur genera rises from 4 at the base of the Campanian to 48 in the upper part. This development is sometimes referred to as the "Campanian Explosion". However, it is not yet clear if the event is artificial, i.e. the low number of genera in the lower Campanian can be caused by a lower preservation chance for fossils in deposits of that age. The generally warm climates and large continental area covered in shallow sea during the Campanian probably favoured the dinosaurs. In the following Maastrichtian stage, the number of North American dinosaur genera found is 30% less than in the upper Campanian.[1]


Animals that lived in the Campanian include:

Cartilaginous fish

[edit]
Cartilaginous fish of the Campanian
Taxa Presence Location Description Images

Chlamydoselachus

Chlamydoselachus

Schizorhiza

Actinopterygii

[edit]
Ray-finned fish of the Campanian
Taxa Presence Location Description Images

Gillicus

100–66 Ma North America A genus of osteoglossomorph fish in the family Ichthyodectidae. Unlike its larger relative, Xiphactinus, by which it was preyed upon, it was a filter feeder instead of a large prey predator.
Gillicus
Ichthyodectes
Xiphactinus


Ichthyodectes

Coniacian to Campanian North America A genus of large osteoglossomorph fish in the family Ichthyodectidae. Like its larger close relative Xiphactinus was a voracious predator.

Xiphactinus

Albian to Maastrichtian North America; Europe; Australia; Venezuela A genus of large osteoglossomorph fish in the family Ichthyodectidae that could reach up to 4 meters in length. Xiphactinus was a voracious predator.

Actinistia

[edit]
Coelacanths of the Campanian
Taxa Presence Location Description Images

Axelrodichthys

  1. Axelrodichthys megadromos
Lower Campanian possibly to early Maastrichtian France A species of averagely sized mawsoniid coelacanth. Related species of the same genus lived mostly on earlier stages of the Cretaceous.
Megalocoelacanthus

Megalocoelacanthus

  1. Megalocoelocanthus dobiei
Lower Campanian possibly to early Maastrichtian USA A very large latimerid coelacanth. It has been estimated to have been 3.5 - 4.5 meters in length.


Dipnoi

[edit]
Lungfish of the Campanian
Taxa Presence Location Description Images

Ceratodus

Olenekian-Eocene Cosmpopolitan A widespread genus of ceratodontiform lungfish in the family Ceratodontidae with multiple species throughout the mesozoic.
Ceratodus
Retodus

Protopterus

Campanian to Recent Egypt, Sudan A ceratodontiform lungfish in the family Protopteridae. Species of this genus, survive to this day.


Retodus

AlbianMaastrichtian Egypt, Sudan A genus of large ceratodontiform lungfish in the family Ceratodontidae.


Lepidosaurs

[edit]
Lepidosaurs of the Campanian
Taxa Presence Location Description Images

Adamisaurus

  1. A. magnidentatus
An agamid iguanian lizard.
Clidastes
Ectenosaurus
Estesia
Globidens alabamaensis
Halisaurus arambourgi
Mosasaurus beaugei
Mosasaurus lemonnieri
Mosasaurus missouriensis
Platecarpus tympaniticus
Prognathodon lutugini
Taniwhasaurus
Telmasaurus
Tylosaurus pembinensis
Tylosaurus proriger

Aiolosaurus

  1. A. oriens
A varanoid lizard of disputed affinity.

Alamitophis

  1. A. argentinus
  2. A. elongatus
A genus of fossil snakes in the extinct family of Madtsoiidae.

Anchaurosaurus

Djadochta Formation, Gobi Desert, Mongolia An iguanian lizard in the extinct clade Gobiguania. Compared to other iguanians, Anchaurosaurus has a relatively elongated skull, large eye sockets, and higher tooth crowns.

Apsgnathus

Brewster County, Aguja Formation, Texas, USA A scincomorph.

Asprosaurus

A genus of anguimorph lizard distantly related to the living Gila monster.

Bainguis

84.9-70.6 mya Bayan Mandahu Formation, Mongolia A diploglossan lizzard.

Catactegenys

Brewster County, Aguja Formation, Texas, USA A large, possibly durophagous, night lizard.

Carusia

A basal anguimorph lizard closely related to the extant Xenosauridae.


Cherminotus

A varanoid lizard that resembles Aiolosaurus.

Clidastes

A relatively small basal mosasaurine marine lizard.

Ctenomastax

Djadochta Formation, Gobi Desert, Mongolia An iguanid iguanian lizard.

Eoxanta

Djadochta Formation, Gobi Desert, Mongolia A scincomorph lizard.


Dryadissector

  1. D. shilleri
A genus of extinct varanoid lizards.

Ectenosaurus

A genus of plioplatecarpine mosasaurs.

Eonatator

  1. E. coellensis
  2. E. sternbergii
A genus of halisaurine mosasaurs.

Estesia

Gobi Desert, Mongolia A genus of anguimorph lizards, originally thought to be closely related to monitor lizards.

Flaviagama

Djadochta Formation, Gobi Desert, Mongolia A priscagamid iguanian lizard.

Globaura

Djadochta Formation, Gobi Desert, Mongolia A scincomorph lizard.

Gobiderma

Djadochta Formation, Gobi Desert, Mongolia An anguimorph lizard that resembled a Gila monster though it was not necessarily closely related to it.

Gobinatus

Djadochta Formation, Gobi Desert, Mongolia A teiid lacertoid lizard.

Globidens

  1. G. alabamaensis
  2. G. dakotensis
  3. G. hisaensis
  4. G. schurmanni
  5. G. simplex
A genus of medium sized mosasaurine mosasaurs. Along with its closest relatives in the tribe Globidensini it had distinctive globular teeth.

Gobiderma

An anguimorph lizard that resembled the modern Heloderma lizards.


Halisaurus

A comparatively small mosasaurs. Type genus of the subfamily Halisaurine.

Herensugea

A madtsoiid snake.

Hymenosaurus

Djadochta Formation, Gobi Desert, Mongolia A scincomorph lizard.

Isodontosaurus

Djadochta Formation, Gobi Desert, Mongolia An iguanian lizard in the extinct clade Gobiguania.

Kawasphenodon

  1. K. expectatus
A sphenodontid rhynchocephalian closely related to the extant tuatara.

Lamarquesaurus

  1. L. cabazai
A basal neosphenodont rhynchocephalian.

Latoplatecarpus

  1. L. nichollsae
  2. L. willistoni
New Zealand, Japan, Antarctica A large plioplatecarpine mosasaur.

Menarana

  1. M. laurasiae
A madtsoiid snake.

Mimeosaurus

A priscagamid iguanian lizard. Mimeosaurus is unique among iguanians in having premaxilla bones at the tip of the snout that are reduced in size, as well as having two pairs of enlarged canine-like teeth in the maxilla

Mosasaurus

  1. M. beaugei
  2. M. conodon
  3. M. lemonnieri
  4. M. missouriensis


A genus of large mosasaurine mosasaurs, contained some of the largest marine reptiles to have ever lived.

Myrmecodaptria

Djadochta Formation, Gobi Desert, Mongolia A basal lizard of contested affinities. Might be related to the contemporary Bainguis.


Odaxosaurus

  1. O. piger
  2. O. priscus
An anguid anguimorph.

Ovoo

  1. O. gurvel'
An early varanid.

Paravaranus

  1. P. angustifrons
A varanoid lizard.

Parmeosaurus

Djadochta Formation, Gobi Desert, Mongolia A scincomorph lizard.


Patagoniophis

  1. P. parvus
A madtsoiid snake.

Phrynosomimus

Djadochta Formation, Gobi Desert, Mongolia A priscagamid iguanian lizard.

Platecarpus

USA, possibly Belgium and Africa A genus of plioplatecarpine mosasaurs.

Pleurodontagama

Bayan Mandahu Formation Gobi Desert, Mongolia Originally thought to be a priscagamid, it's sub-pleurodont dentition instead of acrodont permanent dentition indicates that it might had been a more basal iguanian, a potential "missing link" between Chamaeleontiformes and Pleurodonta.

Plioplatecarpus

  1. P. houzeaui
  2. P. peckensis
  3. P. primaevus
A genus of plioplatecarpine mosasaurs.

Priscagama

Djadochta Formation, Gobi Desert, Mongolia A priscagamid iguanian lizard.

Prognathodon

  1. P. currii
  2. P. lutugini
  3. P. overtoni
A genus of large mosasaurine mosasaurs.

Proplatynotia

  1. P. proplatynotia
An anguimorph lizard closely related to Monstersauria Varanoidea.

Rionegrophis

A madtsoiid snake.

Saichangurvel

Djadochta Formation, Gobi Desert, Mongolia An iguanian lizard in the extinct clade Gobiguania.

Saniwides

A varanoid lizard.

Slavoia

Djadochta Formation, Gobi Desert, Mongolia Previously thought to be a scincomorph, it was later placed in a clade containing Lacertoidea+Scincomorpha. A more recent description of the genus, placed it as a stem-group relative of amphisbaenians within Lacertoidea.


Taniwhasaurus

New Zealand, Japan, Antarctica A medium sized tylosaurine mosasaur.

Temujinia

Djadochta Formation, Gobi Desert, Mongolia An iguanian lizard in the extinct clade Gobiguania.

Telmasaurus

A varanoid lizard.

Tylosaurus

  1. T. bernardi
  2. T. ivoensis
  3. T. pembinensis
  4. T. proriger
  5. T. saskatchewanensis
A genus of large tylosaurine mosasaurs, contained some of the largest marine reptiles to have ever lived.

Zapsosaurus

Djadochta Formation, Gobi Desert, Mongolia An iguanian lizard in the extinct clade Gobiguania.

Testudines

[edit]
Turtles of the Campanian
Taxa Presence Location Description Images

Allopleuron

Archelon
Protostega

Archelon

Atlantochelys

Calcarichelys

Chelosphargis


Ctenochelys

Desmatochelys

Leviathanochelys

Mesodermochelys


Nichollsemys

Pneumatoarthrus

Protostega

Shandongemys

A lindholmemydid pantestudinoid turtle.

Terlinguachelys

Toxochelys

†Plesiosaurs

[edit]
Plesiosaurs of the Campanian
Taxa Presence Location Description Images

Elasmosaurus

80.5 mya Pierre Shale, Kansas, USA Elasmosaurus is a genus of plesiosaur with an extremely long neck.
Elasmosaurus
Styxosaurus

Styxosaurus

83.5-80.5 mya Logan County, Kansas Styxosaurus is an Elasmosaurid plesiosaur.


†Choristoderans

[edit]
Choristoderans of the Campanian
Taxa Presence Location Description Images

Champsosaurus

Campanian to Paleocene North America Champsosaurus is a genus of crocodile-like neochoristodere.
Champsosaurus natator

Crocodylomorphs

[edit]
Crocodylomorphs of the Campanian
Taxa Presence Location Description Images

Acynodon

  1. Acynodon adriaticus
  2. Acynodon iberoccitanus
Early Campanian - Late Maastrichtian 83.5–66 Ma Italy Initially placed within Alligatoridae but has since been reclassified as a more basal globidontan, the oldest and most primitive known to date.
Allodaposuchus precedens
Brachychampsa montana
Deinosuchus riograndensis
Deinosuchus rugosus
Paralligator gradilifrons


Albertochampsa

  1. Albertochampsa langstoni
76.9–75.8 Ma Dinosaur Park Formation, Alberta A basal member of Globidonta, a group of alligatoroids more derived than Deinosuchus or Leidyosuchus that includes living alligators and caimans.

Agaresuchus

  1. Agaresuchus fontisensis
Late Campanian-Maastrichtian Spain A genus of average-sized basal eusuchian. Might be synonymous with its contemporary relative genus Allodaposuchus.

Allodaposuchus

  1. Allodaposuchus precedens
  2. Allodaposuchus iberoarmoricanus
84.9–66.043 Ma Spain and France, A genus of average-sized basal eusuchian, growing to around 3 m (9.8 ft) long. The main feature that distinguishes this species from other related crocodylomorphs is the orientation of a groove at the back of the skull called the cranioquadrate passage; unlike the cranioquadrate passages of other crocodylomorphs, which are only visible at the back of the skull, the cranioquadrate passage of this variety is visible when the skull is viewed from the side. The genus includes many species but it may be paraphyletic in regards to its contemporary relative genera Agaresuchus and Lohuecosuchus.

Artzosuchus

  1. Artzosuchus brachicephalus
Campanian Djadochta Formation, Mongolia Its classification beyond that of a basal crocodylomorph is indeterminant because of the fragmentary nature of the material associated with the genus.

Brachychampsa

  1. Brachychampsa montana
  2. †?Brachychampsa sealeyi
early Campanian to Danian Hell Creek Formation, Montana, USA A globidontan distinguished by an enlarged fourth maxillary tooth in the upper jaw.

Deinosuchus

  1. Deinosuchus hatcheri
  2. Deinosuchus riograndensis
  3. Deinosuchus rugosus
  4. Deinosuchus schwimmeri
82–73 Ma USA A basal alligatoroid and one of the largest crocodylomorphs that ever lived.

Gobiosuchus

  1. Gobiosuchus kielanae
Early Campanian 75–71 Ma Djadokhta Formation, Gobi Desert, Mongolia. A member of Gobiosuchidae, a family of late surviving but basal non-mesoeucrocodilian crocodyliforms.

Leidyosuchus

  1. Leidyosuchus canadensis
Middle Campanian Alberta A basal medium-sized alligatoroid.

Lohuecosuchus

  1. Lohuecosuchus megadontos
Late Campanian-early Maastrichtian Spain A genus of average-sized basal eusuchian. Might be synonymous with its contemporary relative genus Allodaposuchus.

Paralligator

  1. Paralligator gradilifrons
Late Cretaceous, ~96–70 Ma Nemegt Formation, Mongolia A paralligatorid mesoeucrocodylian.

Shamosuchus

  1. Shamosuchus djadochtaensis
83.6–72.1 mya Djadochta Formation, Mongolia A paralligatorid mesoeucrocodylian.

Zaraasuchus

  1. Zaraasuchus shepardi
Zos Canyon, Gobi Desert, Mongolia. A gobiosuchid.

†Pterosaurs

[edit]
Pterosaurs of the Campanian
Taxa Presence Location Description Images

Aerotitan

Campanian-Maastrichtian Allen Formation, Patagonia, Argentina A genus of moderately large azhdarchid pterosaurs.
Aerotitan
Nyctosaurus
Pteranodon

Bogolubovia

Rybushka Formation, Petrovsk, Russia A genus of moderately of medium to large sized azhdarchid pterosaurs.

Geosternbergia

USA, North America Geosternbergia was originally a species of Pteranodon and is famous for its oddly shaped crest.

Montanazhdarcho

Montana, USA Small azhdarchoid pterosaur, probably a tapejarid

Navajodactylus

New Mexico, USA, and Alberta, Canada Known primarily from forearm elements; tentatively assigned to Azhdarchidae, though most likely not part of it.

Nyctosaurus

mid-western United States Nyctosaurus is a genus of pterodactyloid pterosaur.

Piksi

Montana, USA Piksi is a genus of pterosaurs containing the single species Piksi barbarulna.

Pteranodon

Kansas, USA, North America Pteranodon is a genus of pterosaurs which included some of the largest known flying reptiles, with wingspans over 6 metres

Volgadraco

Saratov, Russia A large pterosaur, either an azhdarchoid or a nyctosaurid.

†Dinosaurs

[edit]

†Ornithischia

[edit]
†Ankylosaurs
[edit]
Ankylosaurs of the Campanian
Taxa Presence Location Description Images

Aletopelta

Point Loma Formation, California, USA A medium-sized ankylosaurid, estimated to be around 6 m (20 ft) long.
Edmontonia
Euoplocephalus


Pinacosaurus

Antarctopelta

Santa Marta Formation, James Ross Island, Antarctica A stocky ankylosaur protected by armor plates embedded in the skin. Although a complete skeleton has not been found, the species is estimated to have reached a maximum length of 4 meters (13 feet). Displays characteristics of both ankylosaurids and nodosaurids.

Edmontonia

Campanian to Maastrichtian Horseshoe Canyon Formation, Alberta, Canada A bulky nodosaurid at roughly 6.6 m (22 ft) long. It had small, ridged bony plates on its back and many sharp spikes along its body sides. The four largest spikes jutted out from the shoulders on each side, two of which were split into subspines in some specimens. Its skull had a pear-like shape when viewed from above.

Euoplocephalus


Heishansaurus

A dubious ankylosaurid, possibly a synonym of Pinacosaurus.

Nodocephalosaurus

Palaeoscincus

Judith River Formation known from a single tooth

Panoplosaurus

Judith River Formation, Alberta, Canada; Montana, USA A 5.5–7 m long nodosaurid.

Pinacosaurus

Saichania

Shanxia

Struthiosaurus

Tarchia

Tianzhenosaurus

†Theskelosaurids
[edit]
Parksosaurids of the Campanian
Taxa Presence Location Description Images

Orodromeus

76.7 mya Montana, USA Orodromeus was a small fast bipedal herbivore that probably coexisted with dinosaurs such as Daspletosaurus and Einiosaurus. Its length was estimated by Horner & Weishampel at 2.5 metres.
Orodromeus
†Ceratopsians
[edit]
Ceratopsians of the Campanian
Taxa Presence Location Description Images

Achelousaurus

  1. Achelousaurus horneri
74.2 million years ago Glacier County, Two Medicine Formation, Montana, U.S.A. A centrosaurine.
Achelousaurus
Agujaceratops
Albertaceratops
Anchiceratops
Avaceratops
Bagaceratops
Brachyceratops
Breviceratops
Centrosaurus
Cerasinops
Chasmosaurus
Coahuilaceratops
Coronosaurus
Diabloceratops
Einiosaurus
Graciliceratops
Judiceratops
Kosmoceratops
Lamaceratops
Medusaceratops
Micropachycephalosaurus
Mercuriceratops
Nasutoceratops
Pachyrhinosaurus lakustai
Pentaceratops
Prenoceratops
Protoceratops
Rubeosaurus
Spinops
Styracosaurus
Titanoceratops
Udanoceratops
Utahceratops
Vagaceratops
Xenoceratops
Yehuecauhceratops

Agujaceratops

  1. Agujaceratops mariscalensis
  2. Agujaceratops mavericus
77 million years ago Aguja Formation, Texas A chasmosaurine.

Albertaceratops

  1. Albertaceratops nesmoi

Anchiceratops

  1. Anchiceratops longirostris

Avaceratops

  1. Avaceratops lammersi

Bagaceratops

  1. Bagaceratops rozhdestvenskyi

Bainoceratops

  1. Bainoceratops efremovi

Brachyceratops

  1. Brachyceratops montanensis

Breviceratops

  1. Breviceratops kozlowskii

Centrosaurus

  1. Centrosaurus apertus

Cerasinops

  1. Cerasinops hodgskissi

Ceratops

  1. Ceratops montanus

Chasmosaurus

  1. Chasmosaurus russelli
  1. Chasmosaurus belli

Coahuilaceratops

  1. Coahuilaceratops magnacuerna

Coronosaurus

  1. Coronosaurus brinkmani

Diabloceratops

  1. Diabloceratops eatoni

?Dysganus

  1. Dysganus encaustus
  1. Dysganus bicarinatus
  1. Dysganus peiganus

Einiosaurus

  1. Einiosaurus procurvicornis

Eoceratops

  1. Eoceratops canadensis

Graciliceratops

  1. Graciliceratops mongoliensis

Gryphoceratops

  1. Gryphoceratops morrisoni

Judiceratops

  1. Judiceratops tigris

Kosmoceratops

  1. Kosmoceratops richardsoni

Lamaceratops

  1. Lamaceratops tereschenkoi

Magnirostris

  1. Magnirostris dodsoni

Medusaceratops

  1. Medusaceratops lokii

Mercuriceratops

  1. Mercuriceratops gemini

Micropachycephalosaurus

  1. Micropachycephalosaurus hongtuyanensis

Mojoceratops

  1. Mojoceratops perifania

Monoclonius

  1. Monoclonius crassus

Nasutoceratops

  1. Nasutoceratops titusi

?Notoceratops

  1. Notoceratops bonarellii
Chubut Province, Argentina A dubious genus of possible ceratopsian affinity

Pachyrhinosaurus

  1. Pachyrhinosaurus lakustai


Pentaceratops

  1. Pentaceratops sternbergii

Platyceratops

  1. Platyceratops tatarinovi

Prenoceratops

  1. Prenoceratops pieganensis

Protoceratops

  1. Protoceratops andrewsi
  1. Protoceratops hellenikorhinus

Rubeosaurus

  1. Rubeosaurus ovatus

Spiclypeus

  1. Spiclypeus shipporum

Spinops

  1. Spinops sternbergorum

Styracosaurus

  1. Styracosaurus albertensis

Titanoceratops

  1. Titanoceratops ouranos

Udanoceratops

  1. Udanoceratops tschizhovi

Unescoceratops

  1. Unescoceratops koppelhusae

Utahceratops

  1. Utahceratops gettyi

Vagaceratops

  1. Vagaceratops irvinensis

Xenoceratops

  1. Xenoceratops foremostensis

Yehuecauhceratops

  1. Yehuecauhceratops mudei
72 mya Coahuila, Aguja Formation, Mexico A centrosaurine.


†Pachycephalosaurs
[edit]
Pachycephalosaurs of the Campanian
Taxa Presence Location Description Images

Colepiocephale

Alberta, Canada The oldest known pachycephalosaurid.
Colepiocephale lambei
Goyocephale
Hanssuesia sternbergi
Homalocephale
Prenocephale
Sphaerotholus
Stegoceras
Tylocephale
Wannanosaurus yansiensis

Goyocephale

Mongolia

Gravitholus

Hanssuesia

Alberta, Canada; Montana, USA Distinguished from other pachycephalosaurs by having a depressed parietal region, wide frontoparietal dome, broad nasal characteristics, reduced prefontal lobes, and a reduced parietosquamosal shelf.


Homalocephale

Mongolia Sporting a flat, wedge-shaped skull roof, Homalocephale was different from other pachycephalosaurs.

Ornatotholus

A possible synonym of [Stegoceras]].

Prenocephale

Sphaerotholus

Stegoceras

Tylocephale

Texacephale

77 mya Aguja Formation, Texas

Wannanosaurus

†Ornithopods
[edit]
Ornithopods of the Campanian
Taxa Presence Location Description Images

Acristavus

79 mya Montana and Utah, USA A genus of basal saurolophine hadrosaur.
Acristavus gagslarsoni
Angulomastacator daviesi
Aralosaurus tuberiferus
Brachylophosaurus
Corythosaurus casuarius
Edmontosaurus regalis
Gasparinisaura
Gryposaurus notabilis
Hadrosaurus foulkii
Hypacrosaurus altispinus
Kritosaurus
Lambeosaurus lambei
Malefica
Maiasaura
Naashoibitosaurus ostromi
Nipponosaurus
Ornatops
Parasaurolophus walkeri
Prosaurolophus
Rhabdodon
Saurolophus angustirostris
Shantungosaurus
Tanius
Tsintaosaurus
Velafrons

Angulomastacator

76.9 mya Aguja Formation, Texas A lambeosaurine.

Aquilarhinus

Aguja Formation, Texas Formerly referred to Kritosaurus and Gryposaurus it might be a much more basal hadrosaurid.

Aralosaurus

Upper Santonian-Lower Campanian boundary 85.8 mya Bostobe Formation, Kazakhstan A basal lambeosaurine hadrosaur, Aralosaurus was about the size of an elephant. Although very little is known about Aralosaurus (only one near complete skull has been found); it was identified by a beak with nearly 1,000 small teeth in 30 rows. The back of an Aralosaurus skull was wide, a feature suggestive of large jaw muscles used to power its chewing apparatus.

Brachylophosaurus

76.5 mya Montana, USA; Alberta, Canada Brachylophosaurus was a basal saurolophine hadrosaur which reached an adult length of 9 meters (30 feet).

Corythosaurus

77-76.5 mya Alberta, Canada A derived lambeosaurine. Corythosaurus weighed in at 4 tonnes and measured roughly 10 metres (33 feet) from nose to tail.

Diclonius

75 mya Montana, USA A dubious genus of hadrosaur known only from dental fossils.

Edmontosaurus

73.0-76.5 mya Canada Edmontosaurus, a saurolophine hadrosaur, included some of the largest ornithischian species, measuring up to 12 metres (39 feet) long and weighing around 4.0 metric tons (4.4 short tons).

Gasparinisaura

85 mya Argentina Gasparinisaura was a small ornithopod of the basal clade Elasmaria. In 2010 Gregory S. Paul estimated the length at 1.7 metres, the weight at thirteen kilogrammes.

Gilmoreosaurus

72 mya Mongolia A relatively large hadrosauromorph.

Gryposaurus

83-75.5 mya Alberta, Canada Gryposaurus was a saurolophine hadrosaur of typical size and shape.

Hadrosaurus

79.5 mya New Jersey, USA One of the most basal hadrosaurids and the namesake genus of the family. It was likely bipedal for the purposes of running, but could use its forelegs to support itself while feeding.

Hypacrosaurus

75-67 mya Alberta, Canada A derived lambeosaurine, Hypacrosaurus is most easily distinguished from other hollow-crested duckbills by its tall neural spines and the form of its crest. The neural spines, which project from the top of the vertebrae, are 5 to 7 times the height of the body of their respective vertebrae in the back,[4] which would have given it a tall back in profile. The skull's hollow crest is like that of Corythosaurus, but is more pointed along its top, not as tall, wider side to side, and has a small bony point at the rear

Hypsibema

North Carolina and Missouri, USA A large hadrosauroid known only by a couple of vertebrae.

Kritosaurus

73 mya North America The type specimen of Kritosaurus navajovius is only represented by a partial skull and lower jaws, and associated postcranial remains.

Lambeosaurus

76-75 mya Alberta, Canada A derived lambeosaurine hadrosaur, namesake genus of the subfamily.

Lophorhothon

80 mya Alabama, USA A genus of hadrosauromorphs.

Malefica

Aguja Formation, Texas A basal hadrosaur.

Maiasaura

76.7 mya Montana, USA Maiasaura was a large basal saurolophine hadrosaur, attaining an adult length of about 9 metres (30 feet) and had the typical hadrosaurid flat beak and a thick nose. It had a small, spiky crest in front of its eyes. The crest may have been used in headbutting contests between males during the breeding season.

Microhadrosaurus

China A possibly dubious genus of hadrosaur known by a partial lower jaw from a juvenile.

Mochlodon

Austria A rhabdodontid.

Naashoibitosaurus

73 mya New Mexico, USA Naashoibitosaurus, a saurolophine hadrosaur, based as it is on a single partial skeleton, is not well known in terms of anatomy. Its skull, the most thoroughly described portion, has a low nasal crest that peaks in front of the eyes, but does not strongly arch as in Gryposaurus.

Nipponosaurus

80 mya Sakhalin island A lambeosaurine hadrosaur known from subadult remains.

Ornatops

78 mya New Mexico, USA A genus of basal saurolophine hadrosaur.

Parasaurolophus

76.5-73 mya Alberta, Canada; New Mexico and Utah, USA A large derived lambeosaurine hadrosaur.

Probrachylophosaurus

79 mya Montana, USA; Alberta, Canada A large basal saurolophine hadrosaur which reached an adult length of over 9 meters (30 feet).

Prosaurolophus

76-75 mya Alberta, Canada Prosaurolophus was a large-headed duckbill; the most complete described specimen has a skull around 0.9 meters (3.0 feet) long on a ~8.5 meter long skeleton (~28 ft).[2] It had a small, stout, triangular crest in front of the eyes; the sides of this crest were concave, forming depressions. The upper arm was relatively short.

Pteropelyx

75 mya Montana, USA A dubious genus of lambeosaurine hadrosaurs. Historically, several species were assigned to it, all based on extremely fragmentary remains, but there is no evidence to support these assignments.

Rhabdodon

72 mya France; Spain; Haţeg Island, Romania It is unclear whether it was an iguanodont or a hypsilophodont, and may be a "missing link" between the two. Current evidence indicates it is an iguanodont similar to Tenontosaurus.

Saurolophus

69.5-68.5 mya North America, Asia Saurolophus is known from material including nearly complete skeletons, giving researchers a clear picture of its bony anatomy. S. osborni, the rarer Albertan species, was around 9.8 meters (32 feet) long, with its skull a meter long (3.3 feet). Its weight is estimated at 1.9 tonnes (2.1 tons). S. angustirostris, the Mongolian species, was larger; the type skeleton is roughly 12 meters (39 feet) long, and larger remains are reported.

Shantungosaurus

72 mya China A very large saurolophine hadrosaur closely related to Edmontosaurus, it is one of the longest and largest known hadrosaurids and possibly the largest non-sauropod dinosaur; the composite skeleton of a medium-sized individual mounted at the Geological Institute of China in Beijing measures 14.72 metres (48.3 feet) in length.

Stephanosaurus

76–75 mya Dinosaur Park Formation, Alberta A dubious genus of hadrosaurid dinosaur with a complicated taxonomic history.

Tanius

72–71 mya China A genus of hadrosauromorph dinosaur.

Trachodon

77 mya Montana, USA Trachodon is a dubious genus of hadrosaurid dinosaur based on teeth. Numerous species had been referred to this genus later reassigned to different genera or proven dubious.

Tsintaosaurus

72 mya Southern China A basal but relatively large lambeosaurine hadrosaur.

Velafrons

72 mya Mexico A large derived lambeosaurine.

†Sauropoda

[edit]
Sauropods of the Campanian
Taxa Presence Location Description Images

Alamosaurus

Southwestern United States Alamosaurus is a genus of titanosaurian sauropod dinosaur from the Late Cretaceous Period of what is now North America. It was a large quadrupedal herbivore.
Alamosaurus
Andesaurus
Dreadnoughtus
Huabeisaurus allocotus
Neuquensaurus
Overosaurus
Quaesitosaurus
Saltasaurus

Andesaurus

Neuquén Province, Argentina Andesaurus is a genus of basal titanosaurian sauropod dinosaur.

Argyrosaurus

Lago Colhué Huapi Formation, Argentina

Dreadnoughtus

Cerro Fortaleza Formation, Argentina Dreadnoughtus is one of the largest titanosaurs known.

Gondwanatitan

Adamantina Formation and Cambabe Formation, Brazil

Huabeisaurus

North East, China A member of the Euhelopodidae sauropods.

Laplatasaurus

Allen Formation and Anacleto Formation, both in Argentina; Palacio Formation, Uruguay

Loricosaurus

Campanian-Maastrichtian Allen Formation, Argentina

Microcoelus

Santonian-Campanian Bajo de la Carpa Formation, Argentina

Neuquensaurus

Anacleto Formation, Argentina

Overosaurus

Neuquén Province, Argentina Small-sized titanosaur.

Quaesitosaurus

Shar Tsav, Mongolia Quaesitosaurus is a genus of titanosaurian sauropod.

Saltasaurus

north-west Argentina; Uruguay Saltasaurus is a genus of titanosaurid sauropod dinosaur. An estimated length of 12 metres (39 feet) and a mass of 7 tonnes (8 tons).

Rocasaurus

Campanian-Maastrichtian Allen Formation, Rio Negro Province, Argentina

Theropoda

[edit]

David J. Varrichio observes that during the late Campanian Alberta and Montana had very similar theropods despite significant differences in the types of herbivorous dinosaur faunas.[2]


Non-avian theropods of the Campanian
Taxa Presence Location Description Images

Abelisaurus

Allen Formation?, Anacleto Formation?, Argentina An abelisaurid ceratosaur that probably reached 7 to 9 meters in length; known from only one partial skull.
Abelisaurus
Appalachiosaurus
Archaeornithomimus
Bambiraptor
Carnotaurus
Citipati
Daspletosaurus
Dromaeosaurus
Gobivenator
Mahakala
Oviraptor
Pyroraptor
Saurornithoides
Shuvuuia
Struthiomimus altus
Teratophoneus
Tsaagan
Velociraptor

Appalachiosaurus

A derived tyrannosauroid.

Archaeornithomimus

An ornithomimid ornithomimosaur.

Bambiraptor

A small dromaeosaurid.

Bistahieversor

A large derived tyrannosauroid.

Byronosaurus

A troodontid

Citipati

An oviraptorid oviraptorosaur.

Carnotaurus

A derived abelisaurid ceratosaur that probably reached 7 to 9 meters in length

Chirostenotes

A caenagnathid oviraptorosaur.

Daspletosaurus

  1. Daspletosaurus torosus
  2. Daspletosaurus wilsoni
  3. Daspletosaurus horneri
  4. Daspletosaurus sp.
A genus of large and robustly built tyrannosaurines.

Deinodon

Judith River Formation A dubious tyrannosaurid known from tooth remains.

Dromaeosaurus

A small to medium sized dromaeosaurid maniraptoran theropod with a sickle-like claw.

Dromiceiomimus

An ornithomimid ornithomimosaur.

Dryptosaurus

A derived tyrannosaur.

Dynamoterror

A large basal tyrannosaurine.

Gobivenator

A troodontid

Gorgosaurus

An albertosaurine tyrannosaurid.

Harpymimus

A basal ornithomimosaur.

Khaan

An oviraptorid oviraptorosaur.

Kol

An alvarezsaurid.

Labocania

A large robust theropod of uncertain affinities, possibly a tyrannosaur.

Leptorhynchos

A caenagnathid oviraptorosaur.

Linheraptor

A dromaeosaurid.

Lythronax

Wahweap Formation, Utah A 7-meter tyrannosaurine known from a partially complete skull, some vertebrae and a complete pubis

Luanchuanraptor

A dromaeosaurid.

Mahakala

A small unenlagiid maniraptoran.

Nanshiungosaurus

A relatively large-bodied therizinosaurid.

Noasaurus

A small derived ceratosaur.

Ornithomimus

An ornithomimid ornithomimosaur.

Oviraptor

Mongolia Oviraptor is an oviraptorid oviraptorosaur in the subfamily Oviraptorinae.

Parvicursor

An alvarezsaur.

Pyroraptor

Var, France A small dromaeosaurid.

Richardoestesia

A coelurosaur form taxon known only from teeth and lower jaw fragments.


Saurornithoides

A troodontid.

Saurornitholestes

A dromaeosaurid.

Shuvuuia

An alvarezsaur.

Struthiomimus

An ornithomimid ornithomimosaur.

Teratophoneus

A basal tyrannosaurine.

Thanatotheristes

A close relative of Daspletosaurus.

Troodon

A troodontid.

Tsaagan

A dromaeosaurid.

Variraptor

Var, France A dromaeosaurid.

Velociraptor

Mongolia and China A dromaeosaurid.

Zhuchengtyrannus

Wangshi Group, Zhucheng, China One of the largest tyrannosaurids at between 10–12 meters. Known from a lower jaw and maxilla slightly smaller than those of the later Tyrannosaurus.
Avialae
[edit]
Birds of the Campanian
Taxa Presence Location Description Images

Alexornis

A genus of enantiornithine birds of uncertain affinities.
Gargantuavis
Hesperornis
Hollanda
Ichthyornis


Apatornis

A genus of basal ornithuran birds.

Apsaravis

A genus of basal ornithuromorph birds.

Baptornis

A genus of flightless hesperornithiform birds.

Brodavis

A genus of freshwater hesperornithiform birds. Although Hesperornithes probably lost their flight abilities by the end of the Early Cretaceous, minimal pachyostosis in Brodavis suggests the possibility of some volant abilities.

Canadaga

A genus of flightless hesperornithid birds.

Castignovolucris

An enantiornithe of uncertain affinities. It was estimated to have a wingspan of around 127–185 cm and may have been 75 to 110 cm long when fully grown, making it one of the largest known enantiornitheans to date.

Cretaaviculus

~85 mya Bostobe Formation, Kazakhstan An extinct genus of birds. The type species, C. sarysuensis, is known only from an isolated, asymmetrical contour feather


Elsornis

Elsornis is a genus of basal enantiornithine birds. It was flightless or nearly so.

Gargantuavis

A genus of large, primitive and potentially flightless avialan.

Gettyia

An avisaurid enantiornithe, previously thought to be a species of the later closely related Maastrichtian genus Avisaurus.

Gobipteryx

A genus of euenantiornithine enantiornithe. It possessed a robust beak which convergently evolved with those of modern birds.

Halimornis

A genus of avisaurid enantiornithe. It lived in a coastal environment.

Hesperornis

A genus of flightless hesperornithid birds.

Hollanda

A genus of small ground birds. It was either an enantiornithe or a basal ornithuromorph.

Ichthyornis

A genus of toothy seabird-like ornithuran.

Intiornis

A genus of avisaurid enantiornithe. Although closely related to some of the largest avisaurids, members of this genus were very small birds.

Maaqwi

A genus of large marine diving vegaviid bird with an estimated body mass of 1.5 kg.

Mirarce

A genus of avisaurid enantiornithes, closely related to the later genus Avisaurus.

Limenavis

Limenavis is the closest known relative of modern birds.

Neogaeornis wetzeli

A marine bird from Chile. It had the midfeet of a foot-propelled diving bird, but its relationships are enigmatic. The only known species is from the Campanian-Maastrichtian boundary.

Yandangornis

A genus of basal long-tailed avialan. May potentially be a non-avialan genus of theropod.

Mammals

[edit]
Mammals of the Campanian
Taxa Presence Location Description Images

Alphadon

A basal metatherian.

Didelphodon

A basal metatherian.

Kamptobaatar

A multituberculate allotherian.

Kennalestes

An insectivorous leptictid eutherian.

Kryptobaatar

Also known as Gobibaatar[3] or Tugrigbaatar[4]. It's a multituberculate allotherian.

Zalambdalestes

Type genus of Zalambdalestidae, a basal clade of Asian eutherians.
  1. ^ See Weishampel et al. (2004)
  2. ^ Cite error: The named reference mt-ovi-abs-42 was invoked but never defined (see the help page).
  3. ^ Kielan-Jaworowska 1970, pp. 35–49.
  4. ^ "Gobibaatar has page priority but was selected as the junior synonym by Kielan-Jaworowska in 1980," (McKenna & Bell, 1997)